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Paul Giamatti as Story Lord, conductor of the Story Train.

Eddie Pepitone as Testicle Monster #2, a Time Cop from the fourth dimension.

Keegan-Michael Key as Testicle Monster #1, a Time Cop from the fourth dimension.

Liam Cunningham as Balthromaw, a dragon who inadvertently soul bonds with Rick.

Matthew Broderick as Talking Cat, an alien cat who communicates through telepathy.

Justin Theroux as Miles Knightly, a master criminal and organiser of HeistCon.

Elon Musk as Elon Tusk, the CEO of Tuskla.

Adlon also voices Angie Flynt, a retired criminal.

Pamela Adlon as Vermigurber's Child, whose life Rick threatens when attempting to track down Tony.Jeffrey Wright as Tony, a kindhearted yet depressed alien salaryman who accidentally crosses Rick.Taika Waititi as Glootie, Rick's intern and a member of the Monogatron species.Kathleen Turner as Monogatron Queen, who loves podcasts and fights with her husband.Sam Neill as Monogatron Leader, the leader of the species who attempts to conquer Earth.

Sherri Shepherd as the Judge, a judge who oversees Morty's trial.Sarah Chalke as Beth Smith, Rick's daughter, and Morty's and Summer mother a generally level-headed person, who is dissatisfied with her marriage.Spencer Grammer as Summer Smith, Rick's granddaughter, and Morty's sister a conventional teenager who worries about improving her status among her peers.Chris Parnell as Jerry Smith, Rick's son-in-law, and Morty's and Summer's father a simple-minded and insecure person, who disapproves of Rick's influence over his family.Justin Roiland as Rick Sanchez and Morty Smith, the two main characters of the show Rick is an eccentric mad scientist and Morty is his kind but easily distressed grandson.See also: List of Rick and Morty characters Main cast

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